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\frac{d}{d t} (q_1) = - v_r \frac{d}{d t} (q_2) = (- 2) v_r \frac{d}{d t} (q_3) = v_r \frac{d}{d t} (q_4) = 2 v_r u_f = u_1 + 2 u_2 u_r = u_3 + 2 u_4 u_1 = RT \ln (K_q_1 q_1) u_2 = RT \ln (K_q_2 q_2) u_3 = RT \ln (K_q_3 q_3) u_4 = RT \ln (K_q_4 q_4) v_r = kappa_r (ⅇ^{\frac{u_f}{RT}} - (ⅇ^{\frac{u_r}{RT}}))

Source: Derived from workspace The Physics of Physiology - Example 5: A biochemical reaction with mixed stoichiometry at changeset 385e62f15780.

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FAIRDO BG example 3.5.cellml